About the Centers
Basic Science Trainees
The Iowa Wellstone Center originated with a collaboration among the Center investigators, which pre-dates NIH funding. In 1997, Drs. Campbell, Moore and Mathews began regular meetings to review biopsies. Basic science trainees in the Campbell Laboratory joined these meetings, with discussions about biopsy findings and unusual clinical cases often sparking new research directions. With Dr. Campbell’s mentorship, research conducted by graduate student and postdoctoral trainees is making a significant contribution to our understanding of the molecular pathogenesis of diverse muscular dystrophies.
Prospective applicants should send a letter of application, curriculum vitae, and names of three references to Dr. Kevin Campbell.
![Dr. Kevin Campbell photo](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Campbell_Kevin.jpg?h=7d929dff&itok=xBT94fuh)
Kevin P. Campbell, PhD
Title/Position
Director, Senator Paul D. Wellstone Muscular Dystrophy Specialized Research Center
Investigator, Howard Hughes Medical Institute
Professor and Chair, Department of Molecular Physiology and Biophysics
Roy J. Carver Biomedical Research Chair in Molecular Physiology and Biophysics
Professor of Neurology
Graduate Student Trainees
Current
![emma luhmann](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2024-03/Emma%20Luhmann.jpg?h=d07df597&itok=z14adlG6)
Emma Luhmann
Title/Position
Graduate Student Trainee (2024-)
Former
![Ameya Walimbe](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Ameya%20Walimbe.jpg?h=bf372811&itok=X2OVXdSk)
Ameya Walimbe, MD, PhD
Title/Position
Graduate Student Trainee (2012-2020)
Child Neurology Fellow, Baylor College of Medicine, Houston, TX
![Jakob Satz](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Jakob Satz, PhD
Title/Position
Graduate Student Trainee (1999-2007)
Postdoctoral Fellow, Susan L. Ackerman Lab, HHMI, The Jackson Laboratory, Bar Harbor, ME
![Dimple Bansal](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Dimple%20Bansal.jpg?h=715cc7c5&itok=1Si7KA72)
Dimple Bansal, PhD
Title/Position
Graduate Student Trainee (1998-2003)
Independent Entrepreneur, Drug Discovery Oncology & DMD, Pre-clinical drug development Oncology, Marketing & Business Development, McLean, Virginia
![Myoung-Goo Kang](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Myoung-Goo%20Kang.jpg?h=da692b23&itok=GTVFiXZ7)
Myoung-Goo Kang, PhD
Title/Position
Graduate Student Trainee (1998-2003)
Staff Scientist, Baylor College of Medicine, Houston, TX
![Jyoti Arikkath](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Jyoti%20Arikkath_0.png?h=9914e63a&itok=h2E_IfxX)
Jyothi Arikkath, PhD
Title/Position
Graduate Student Trainee (1997-2002)
Scientific Review Officer, Division of Neuroscience, Development and Aging, Center for Scientific Review, NIH, Bethesda, MD
![Leland Lim](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Leland Lim, MD, PhD
Title/Position
Graduate Student Trainee (1993-1999)
Clinical Associate Professor, Department of Neurology and Neurological Sciences, Stanford University School of Medicine, Stanford, CA
![Chris Gurnett.](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Chris%20Gurnett.jpg?h=bf372811&itok=Joy4KzoQ)
Christina (Chris) Gurnett, MD, PhD
Title/Position
Graduate Student Trainee (1992-1997)
A. Ernest and Jane G. Stein Professor of Neurology and Director, Division of Pediatric and Developmental Neurology, Washington University School of Medicine, St. Louis, MO
Postdoctoral Trainees
Current
![Miguel Gonzales](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Miguel%20Gonzales%20v2.jpg?h=35b140b0&itok=XM4u-b-G)
Miguel F. Gonzales, PhD
Title/Position
Postdoctoral Scholar (2022-Present)
![Divya Rathi](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Divya%20Rathi%203.jpg?h=bf372811&itok=pzdhMg90)
Divya Rathi, PhD
Title/Position
Postdoctoral Scholar (2022-Present)
![Ishita Chandel](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Ishita%20Chandel.png?h=5fc361b6&itok=mdsu_reK)
Ishita Chandel, PhD
Title/Position
Postdoctoral Scholar (2021-Present)
![Jeff Hord](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Jeff%20Hord.jpg?h=bf372811&itok=x-56D1oX)
![Soumya Joseph](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Soumya%20Joseph.jpg?h=bf372811&itok=kE_7b66K)
Former
![Hide Okuma](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Hide%20Okuma.jpg?h=bf372811&itok=d73cumgx)
Hidehiko Okuma, MD, PhD
Title/Position
Postdoctoral Scholar (2016-2021)
Anesthesiologist, Kameda Medical Center, Kamogawa City, Japan
![Tiandi Yang](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Tiandi%20Yang.jpg?h=bf372811&itok=D4QeA2O0)
Tiandi Yang, PhD
Title/Position
Postdoctoral Scholar (2017-2020)
Postdoctoral Fellow, Department of Immunology, Harvard Medical School, Cambridge, MA
![Toru Egashira](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Toru%20Egashira_0.png?h=d07e2d79&itok=CiXGk1Fq)
Toru Egashira, MD, PhD
Title/Position
Postdoctoral Scholar (2014-2019)
Chief Physician, Department of Cardiology, Tachikawa Hospital, Tokyo, Japan
![Takahiro Yonekawa](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Takahiro%20Yonekawa.png?h=70d6812a&itok=U2YAFXhU)
Takahiro Yonekawa, MD, PhD
Title/Position
Postdoctoral Scholar (2014-2018)
Assistant Professor, Department of Pediatrics, Mie University Graduate School of Medicine, Mie, Japan
![Tianqing Zheng](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Tianqing%20Zheng.png?h=f765fa0c&itok=7S9YO-Te)
Tianqing Zheng, PhD
Title/Position
Postdoctoral Scholar (2014-2015)
Postdoctoral Researcher, Lerner Lab, The Scripps Research Institute, San Diego, CA
![Jessica de Greef](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Jessica%20de%20Greef.jpg?h=bf372811&itok=QhH2fhzV)
Jessica de Greef, PhD
Title/Position
Postdoctoral Scholar (2010-2015)
Assistant Professor, Department of Human Genetics, Leiden University Medical Center, The Netherlands
![Jennifer Levy](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Jennifer%20Levy_2.png?h=b68768d0&itok=hvHm72--)
Jennifer Levy, PhD
Title/Position
Postdoctoral Scholar (2008-2016)
Scientific Director, Coalition to Cure Calpain 3 (C3), Bryn Mawr, PA
![Matthew Goddeeris](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Matthew%20Goddeeris.png?h=4286dde7&itok=MkUKFmDJ)
Matthew Goddeeris, PhD
Title/Position
Postdoctoral Scholar (2007-2014)
Vice President of Biology, Cellarity, Cambridge, MA
![Kei-ichiro Inamori](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Kei-ichiro Inamori, PhD
Title/Position
Postdoctoral Scholar (2006-2013)
Associate Professor, Institute of Molecular Biomembrane and Glycobiology, Tohoku Medical and Pharmaceutical University, Sendai, Japan
![Erik Rader](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Erik Rader, PhD
Title/Position
Postdoctoral Scholar (2006-2012)
Senior Service Fellow, National Institute for Occupational Safety and Health, Centers for Disease Control and Prevention, Morgantown, WV
![Rolf Turk](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Rolf%20Turk.png?h=dad9a325&itok=vPJKjDkS)
Rolf Turk, PhD
Title/Position
Postdoctoral Scholar (2005-2015)
Research Scientist, Integrated DNA Technologies, Coralville, IA
![Yuji Hara](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Yuji%20Hara.png?h=48f646f4&itok=6PjRjtz1)
Yuji Hara, PhD
Title/Position
Postdoctoral Scholar (2005-2012)
Professor, Department of Pharmaceutical Sciences, University of Shizuoka, Japan
![Tobias Willer](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Tobias%20Willer.png?h=54c7d562&itok=wYwjbaTf)
Tobias Willer, PhD
Title/Position
Postdoctoral Scholar (2004-2009)
Director, Discovery Research and Gene Therapy Science, Amicus Therapeutics - Gene Therapy Center of Excellence
![Daniel Beltran](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Daniel%20Beltran.png?h=89c8fbe6&itok=_2GIYtB_)
Daniel Beltran, PhD
Title/Position
Postdoctoral Scholar (2004-2008)
Lead of Metabolic Operations, Clinical Genomics, Invitae, San Francisco, CA
![Renzhi Han](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Renzhi%20Han.png?h=a690c668&itok=llF_2hn4)
Renzhi Han, PhD
Title/Position
Postdoctoral Scholar (2003-2009)
Professor of Pediatrics, Indiana University School of Medicine, Indianapolis, IN
![Kazuhiro Kobuke](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Kazuhiro Kobuke, MD, PhD
Title/Position
Postdoctoral Scholar (2003-2009)
Manager, Division of Cardiology, Shimada Municipal Hospital, Shimada City, Shizuoka, Japan
![Aaron Beedle](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Aaron%20Beedle.png?h=ee1ba6e7&itok=TUZ31Jq3)
Aaron Beedle, PhD
Title/Position
Postdoctoral Scholar (2003-2008)
Department Chair and Associate Professor, Department of Pharmaceutical Sciences, SUNY Binghamton University, Binghamton, NY
![Takako Yoshida-Moriguchi](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Takako%20Yoshida-Moriguchi.png?h=b210566b&itok=83jxg-K_)
Takako Yoshida-Moriguchi, PhD
Title/Position
Postdoctoral Scholar (2003-2008)
Director, Metabolic Diseases, Sigilon Therapeutics, Inc., Cambridge, MA
![Yoshiaki Saito](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Yoshiaki Saito, MD, PhD
Title/Position
Postdoctoral Scholar (2003-2004)
Chief of Inpatient Ward, Department of Child Neurology, National Center Hospital of Neurology and Psychiatry, Tokyo, Japan
![Motoi Kanagawa](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Motoi%20Kanagawa.png?h=5bc8c916&itok=XPMkYkyu)
Motoi Kanagawa, PhD
Title/Position
Postdoctoral Scholar (2001-2005)
Professor, Department of Molecular Cardiovascular Biology and Pharmacology, Ehime University Graduate School of Medicine, Ehime, Japan
![Severine Groh](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Severine Groh, PhD
Title/Position
Postdoctoral Scholar (2001-2008)
Associate Professor, Division of Biological and Physical Sciences, Delta State University, Cleveland, MS
![Yvonne Kobayashi](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Yvonne%20Kobayashi.png?h=05d3f490&itok=YIq-KbR6)
Yvonne Kobayashi, PhD
Title/Position
Postdoctoral Scholar (2000-2008)
Senior Director, Project Management and Partner Relationships, Prevail Therapeutics, New York, NY
![Hajime Kusano](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Hajime Kusano, PhD
Title/Position
Postdoctoral Scholar (2000-2005)
Manager, Business Development & Licensing, Tokyo, Japan
![Dan Michele](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Dan%20Michele%20photo_1.jpg?h=03d48e0f&itok=vjD8hxIZ)
![Fumiaki Saito.](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Fumiaki%20Saito.png?h=ae654baa&itok=-b1bGg8-)
Fumiaki Saito, PhD
Title/Position
Postdoctoral Scholar (1999-2002)
Associate Professor, Department of Neurology, Teikyo University School of Medicine, Tokyo, Japan
![Rita Barresi](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Rita%20Barresi.png?h=1fd42b47&itok=RUBhSsyT)
Rita Barresi, PhD
Title/Position
Postdoctoral Scholar (1998-2006)
Head of Neurobiology Lab, San Camillo IRCCS SRL, Venice, Italy
![Chien-Chang Chen](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Chien-Chang%20Chen.jpg?h=bf372811&itok=5y9Z6E5-)
Chien-Chang Chen, PhD
Title/Position
Postdoctoral Scholar (1998-2004)
Research Fellow, Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan
![Federica Piccolo](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Federica%20Piccolo.jpg?h=62dbf364&itok=HgY603Ab)
Federica Piccolo, PhD
Title/Position
Postdoctoral Scholar (1998-2002)
Funding Programme Manager, Guido Berlucchi Foundation, Brescia, Italy
![Ronald Cohn](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Ronald%20Cohn.jpg?h=bf372811&itok=12egKXsF)
Ronald Cohn, MD, PhD
Title/Position
Postdoctoral Scholar (1998-2001)
President & CEO, The Hospital for Sick Children, Toronto, Ontario, Canada
![Madeleine Durbeej-Hjalt](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
![Connie Lebakken](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Connie Lebakken, PhD
Title/Position
Postdoctoral Scholar (1997-2001)
President and Chief Operating Officer, Stem Pharm, Inc., Madison, WI
![Ramon Coral-Vazquez](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Ramon Coral-Vazquez, PhD
Title/Position
Postdoctoral Scholar (1996-1999)
Senior Researcher, Instituto Politécnico Nacional, Mexico City, Mexico
![Hiroki Yamada](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Hiroki Yamada, MD, PhD
Title/Position
Postdoctoral Scholar (1996-1999)
Division Chief, Division of Neurology, Tokyo Metropolitan Hiroo Hospital, Tokyo, Japan
![Kathleen Holt](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Kathleen Holt
Title/Position
Postdoctoral Scholar (1995-2000)
Senior Director, Eukaryotic Research, Locus Biosciences, Inc., Morrisville, NC
![Rachelle Crosbie-Watson](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Rachelle%20Crosbie-Watson.png?h=473d1f98&itok=gH2RR-HC)
Rachelle Crosbie-Watson, PhD
Title/Position
Postdoctoral Scholar (1995-1999)
Professor and Chair, Department of Integrative Biology and Physiology, UCLA, Los Angeles, CA
![Michael Henry](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Michael%20Henry.png?h=68265b8b&itok=MO_uTdEl)
Michael Henry, PhD
Title/Position
Postdoctoral Scholar (1995-1999)
Interim Director, Holden Comprehensive Cancer Center and Professor, Department of Molecular Physiology and Biophysics, University of Iowa, Iowa City, IA
![Ricardo Felix](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Ricardo Felix, PhD
Title/Position
Postdoctoral Scholar (1995-1998)
Professor, Department of Cell Biology, Center for Research and Advanced Studies of the National Polytechnic Institute (Cinvestav-IPN), Mexico City, Mexico
![Volker Straub](/sites/wellstone.medicine.uiowa.edu/files/styles/square__768_x_768/public/2023-12/Volker%20Straub.png?h=68265b8b&itok=an07cqHw)
Volker Straub, MD, PhD
Title/Position
Postdoctoral Scholar (1995-1998)
Harold Macmillan Professor of Medicine, Institute of Human Genetics, Newcastle University, Newcastle upon Tyne, UK
![Franck Duclos](/sites/wellstone.medicine.uiowa.edu/files/styles/square__1024_x_1024/public/externals/4/a/4aef231b1948f3e4b39544f3515e4493.png?itok=a8jd4cGl)
Franck Duclos, PhD
Title/Position
Postdoctoral Scholar (1994-1999)
Senior Principal Scientist, Cardiovascular & Fibrosis Discovery Biology, Bristol-Myers Squibb, Pennington, NJ